Y.-L. Zhou, J.A. Nicholls, Z.-H. Liu, D. Hartley, A. Szito, A. Ślipiński and A. Zwick (2022) Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma granarium Everts. Insect Systematics and Diversity, 6(6), 2; 1–24.
1. Subfamily Orphilinae LeConte, 1861
Orphilus Erichson, 1846
Ranolus Blair, 1929
2. Subfamily Trinodinae Casey, 1900
2.1 Tribe Trinodini (= Trinoparvini syn. nov.)
Apsectus LeConte, 1854
Evorinea Beal, 1961
Trinodes Dejean, 1821
Trinoparvus Háva, 2004
2.2 Tribe Thylodriini
Thylodrias Motschulsky, 1839
2.3 Tribe Trichelodini
Hexanodes Blair, 1941
Trichelodes Carter, 1935
Trichodryas Lawrence & Ślipiński, 2005
3. Subfamily Dermestinae Latreille, 1804
3.1 Tribe Dermestini
Dermestes Linnaeus, 1758
Derbyana Lawrence & Ślipiński, 2005 stat. nov.
Dermestinus Zhantiev, 1967
Dermalius Háva, 2001
Montandonia Jacquet, 1886
3.2 Tribe Marioutini
Mariouta Pic, 1898
Rhopalosilpha Arrow, 1929 monotypic
3.3 Tribe Thorictini stat. nov. (=Thorictinae syn. nov.)
Afrothorictus Andreae, 1967
Macrothorictus Andreae, 1967
Thorictodes Reitter, 1875
Thorictus Germar, 1834
4. Subfamily Trogoparvinae subfam. nov.
Trogoparvus Háva, 2001
5. Subfamily Attageninae Laporte de Castelnau, 1840
5.1 Tribe Adelaidiini
Adelaidia Blackburn, 1891
5.2 Tribe Attagenini
Novelsis Casey, 1900
Lanorus Mulsant & Rey, 1868 rev. nov. (= Paranovelsis Casey, 1900 syn. nov.)
Attagenus Latreille, 1802
Aethriostoma Motschulsky, 1858 rev. nov.
Katkaenus Háva, 2006
Telopes Redtenbacher, 1843 rev. nov.
▲Genera of uncertain status:
Adelaidella Zhou et al., 2020
Araphonotos Beal & Kadej, 2008 monotypic
Apphianus Beal, 2005 monotypic
Egidyella Reitter, 1899
Sefrania Pic, 1899 monotypic
6. Subfamily Megatominae Leach, 1815
6.1 Tribe Anthrenini (=Dermeanthrenini)
Anthrenus Geoffroy, 1762
Dermeanthrenus Háva, 2008 monotypic
6.2 Tribe Ctesiini
Ctesias Stephens, 1830
6.3 Tribe Megatomini
6.3.1 Subtribe Megatomina
Globicornis Latreille, 1829
Dearthrus LeConte, 1861
Hirtomegatoma Pic, 1931
Megatoma Herbst, 1791
Zhantievus Beal, 1992
6.3.2 Subtribe Orphinina subtribe nov.
Caccoleptus Sharp, 1902
Curtophinus Pic, 1954 rev. nov.
Falsoorphinus Pic, 1931 rev. nov.
Labrocerus Sharp, 1885
Orphinus Motschulsky, 1858 (= Jiriella Kitano, 2013 syn.nov.)
Thaumaglossa Redtenbacher, 1867
Zahradnikia Háva, 2013
6.3.3 Subtribe Trogodermina
Claviella Kalik, 1987 monotypic
Cryptorhopalum Guérin-Méneville, 1838 (= Hemirhopalum Sharp, 1902 syn. nov.)
Eurhopalus Solier in Gay, 1849 (= Sodaliatoma Háva, 2013 syn. nov., Reesa Beal, 1967 syn. nov., Neoanthrenus Armstrong, 1941 syn. nov., Anthrenocerus Arrow, 1915
syn. nov., Myrmeanthrenus Armstrong, 1945 syn. nov.)
Phradonoma Jacquelin du Val, 1859
Trogoderma Latreille, 1821
▲Genera of uncertain status:
Caccoleptoides Herrmann, Háva & Kadej, 2015 monotypic
Paratrogoderma Scott, 1926
Turcicornis Háva, 2000 monotypic
Valdesetosum Háva, 2015 monotypic
Volvicornis Háva & Kalík, 2004 monotypic
