S Shibuya, K Kiritani and Kenji Fukuda (2020) Life history traits of five ground beetle (Coleoptera: Carabidae) species common in Honshu Island, Japan. Acta Zoologica Academiae Scientiarum Hungaricae, 66 (Suppl.): 185–198.
5種Harpalus griseus, H. eous, H. tridens, Synuchus cycloderus and Carabus procerulusについて飛翔筋の発達などを見ている。