Vasilikopoulos et al. (2020) An integrative phylogenomic approach to elucidate the evolutionary history and divergence times of Neuropterida (Insecta: Holometabola). BMC Evolutionary Biology, 20: 64. https://doi.org/10.1186/s12862-020-01631-6
系統樹を見るとツノトンボ科を認めているようだ。？？
