M.J. Paulsen (2018) Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini). Insecta Mundi 0624: 1–13.
ツノヒョウタンクワガタ属のobesus種群を再検討し、Nigidius gravelyi Paulsenをボルネオから記載。
M.J. Paulsen (2018) Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini). Insecta Mundi 0624: 1–13.
ツノヒョウタンクワガタ属のobesus種群を再検討し、Nigidius gravelyi Paulsenをボルネオから記載。